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Digizz, no expert on fish tanks or material strenghts but I think you need to over engineer the construction of the cabinet. Perhaps a torsion box platform to support the tank and either a steel or hardwood frame to in turn strengthen the whole base of the cabinet. At least this way you could use the MDF or veneered ply as a decorative element and hence only need to use much thinner stock and also you could lose the centre support. Just a thought or two but with the tremendous weight I'd play safe.
Noel
Noel